11/09/01
Name of disorder Early Embryonic Death (EED) Species of animal affected Although
EED can
happen in any species the paper will deal with it in the horse. In horses, EED is thought to be the cause of
infertility in nearly twenty percent of the mares and higher than seventy
percent in subfertile mares. EED is
considered loss of the embryo before the fortieth day of gestation. Many mares show little or no signs of EED,
there is little to no vaginal bleeding after EED. Some noticeable signs of EED include failure to return to heat at
the normal time and if the EED happens after the thirty-sixth day of gestation
the endometrail cups are already functional and will secrete chorionic
gonadotropin for 110 to 120 days even with out the presence of a fetus. This
will keep the mare from showing signs of heat and may delay her breeding to the
next season. Causative agent EED can
be caused by many factors including chronic endometritis, progesterone
deficiency or chromosomal defects.
Chronic endometritis is an infection of the lining of the uterus caused
commonly by bacteria such as streptococcus zooepidemicus, E. coli, Pseudomonas
aeroginosa can cause EED through three different paths. The first path occurs
before the seventeenth day of gestation, when the embryo is free moving and is
nourished by secretions for the endometrail glands. If these glands do not produce enough secretions to support the
embryo during this phase the embryo will not survive. The second pathway for chronic endometritis to cause EED involves
release of prostaglandin from infected endometrium. This release of hormones
causes the corpus luteum to lysis prematurely, causing progesterone levels,
necessary for maintaining pregnancy, to drop and the termination of the
pregnancy. The third pathway is caused of the bacteria and inflammation
products as a result of the infection of the endometrium. The products and effects of these bacteria
can prove lethal for embryos during the early stages of pregnancy. Low
levels or varying levels of progesterone during the pregnancy can also pose a
treat to embryos. With out consistent and high levels of progesterone the
mare’s body cannot sustain a pregnancy and her body may abort the conceptous in
favor of returning to estrus. This variation in the levels of progesterone can
because by chronic endometritis as well as other factors. While chromosomal defects in horses are much
less common in horses then in humans, many of these defects are incompatible
with early embryonic development and cause death of the embryo. Prevention and Control Preventing
EED by diagnosing choric endometritis before breeding, using an endometrail
culture and biopsy, and treating the endometritis can reduce cases of EED and
improve fertility in a number of mares. Treating endometritis can involve
antibiotics to treat the bacterial infection and or flushing of the uterus to
remove and clean the infection and inflammation. Endometritis can also be prevented with proper hygienic practices
and good conformation in mares. Reducing the number of services per mare and
using artificial breeding technologies where possible help to reduce the change
of infection with in the uterus. Some
conformational faults can be surgically corrected like pneumovagina, urovagina
and incompetent cervix, all of which make a mare at higher risk for chronic
endometritis. Ultra
sounding infertile mares to follow the progress of embryonic development can
help to find what is causing the EED by determining when the embryo is
dying. If the embryo is surviving past
day eight but then dies in the following days embryo transfer can be used to
produce a foal from a mare and avoid EED. Embryo transfer techniques are used
to remove and embryo between day six and day eight post ovulation. By
transferring the embryo to a healthy recipient mare the donor mare’s
reproductive problems can be bypassed to produce a foal with her genetics. Blood
test to determine the concentration of blood progesterone is a good indicator
of weather of not the mare is producing high and consistent progesterone
levels. Unstable levels of progesterone
can be corrected by administering regumate, an oral progesterone to the mare
for the first 180 days of pregnancy, this will reduce the variation in the
hormone level and prevent EED. After
the 180-day treatment period the placenta takes over the function of
maintaining hormone levels and the oral progesterone is no longer needed. Darling, Kjersten D.V.M, and James M. Griffin, M.D. Veterinary
Guide to Horse Breeding. New York: Howell Book House, 1999. Georgi, R. P., Et al. The Horse.
2nd ed. New York: W. H. Freeman and Company, 1990 Squires, E.L., P.M. McCue and D. Vanderwall. “The
Current Status of Equine Embryo Transfer.” Theriogenology 51 (1999):
91-104.
Effects
on reproduction
Work Cited